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Furthermore it raises the issue of the presence of unnoticed environmental effects that may inflate estimates of heritability. Hence, due to difficulties in detecting genetic variance and covariances and the potential influenceof environment dependent parental effects on heritability, our ability to predict evolutionarychange in highly plastic traits may be limited. Interestingly, such changes in the environment are not necessarily constant over time as they often show high levels of fluctuation. However,selection in a fluctuating environment can alter sex-specific patterns of genetic and environmentalvariation, indicating support for the adaptive canalization hypothesis. Interestingly we found substantial sex-specificgenetic effects and low genetic correlations between sexes r MF in traits that are only moderately weight at emergence to slightly longevity sexually dimorphic. Here we use a quantitativegenetic framework to test these hypotheses and to investigate both immediate changes in, and theevolution of, sexual dimorphism in response to a changing environment with and withoutfluctuations. This is for instance characterized by an increase in temperature.

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Here we address the question of whether the relativeimportance of these mechanisms is related to the degree of trait plasticity. Uppsala universitet, Teknisk-naturvetenskapliga vetenskapsområdet, Biologiska sektionen, Institutionen för ekologi och genetik, Zooekologi Upphörd Our results suggest that fluctuations in the environment have major impact on the response of a population to environmental change; in a highly variable environment with low predictability, a plastic response might not be beneficial and the response is genetically and environmentally canalized resulting in a low potential to respond to selection and low environmental sensitivity. Björklund, Mats Uppsala universitet, Teknisk-naturvetenskapliga vetenskapsområdet, Biologiska sektionen, Institutionen för ekologi och genetik, Zooekologi. We investigate theinfluence of additive genetic effects and non genetic parental effects in traits that differ in theirdegree of plasticity. Fourth, I concentrated on sex differences in response to temperature, and finally, I assessed the relative importance of genetic and nongenetic inheritance for traits that differ in their plastic response to a change in the environment.

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